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The 3rd Moscow International Conference program features contributions in the broad area of molecular evolution and its applications: ancient genetic history of life, algoritms and methodology to analyze genetic data, evolutionary genomics, phylogenomics, fundamental and applied phylogenetics. The Moscow third conference on Molecular Phylogenetics (MolPhy) is organized jointly by the Science and Education Centers "Evolutionary Genomics and Bioinformatics" (Moscow State University: Belozersky Institute for Physicochemical Biology, Faculty of Biology, Faculty of Bioengineering and Bioinformatics) and "Bioinformatic and Computtational Technologies"(the Institute for Information Transmission problems, Russian Academy of Sciences) with support by leading academic organizations ans scientific funds.