Аннотация:HYPOSTASE–‘CHALAZAL INCLUSION’ FUNCTIONAL COMPLEX IN CORE CARYOPHYLLALES AND METABOLITE TRANSPORT IN THE SEED. Hypostase of core Caryophyllales is a compound unit. The core hypostase originates from the nucellus–chalaza boundary tissue and incorporates adjoining cells of the nucellus in all 4 species concerned. It also incorporates some underlaying cells of the chalaza in Lychnis coronaria. Group of chalaza cells under the hypostase differentiates into the so called ‘chalazal inclusion’ of small thin-walled cells with large nucleus. The hypostase cells are more durable than they were previously thought. Their lifespan differs in different species investigated, but it is invariably much shorter than the period of time the seed takes to develop. These cells continue to transport assimilates for a while after they have died off. The lifespan of the ‘chalazal inclusion’ cells exceeds that of their hypostase counterparts. They are still alive in a seed that has accomplished its morphogenesis. The ‘chalazal inclusion’ cells show some haustorium properties. The ‘chalazal inclusion’ cells seem to cooperate with the hypostase ones and then substitute the latter in transporting assimilates.